Hypecoum.] FLORA INDICA. 275
questioned. The same argument we consider opposed to Lindley's view of the sepals
of Fumariacea being bracts, the outer petals, sepals, etc., the inner alone true pe-
tals, chiefly because the relation of the stamens to the petals is, as above stated, ex-
actly what occurs in the Berberideous genera mentioned above, and because Meoonia
in Papaveracea has also only four petals and as many stamens.
De Candolle (Syst. ii. 101) says, that according to Sehultes, Obs. Bot. 26, H. pro-
cumbens has sometimes four sepals and six stamens. This increase of the number of
sepals is further opposed to the view of the sepals being bracts, and the presence of
six stamens reduces the genus to the hexandrous type common to all other Fumari-
acea. The position of the two additional sepals and stamens is not given by De
Candolle; but if the six latter consist of two pairs, opposite to the smaller petals, and
consequently to the outer sepals, it affords an additional point of affinity between
Pumariacea and Cmcifera, and favours M« Gray's views of the identity of the sta-
minal formula in these two Orders (Ann. Sc. Nat. ser, 2. xviiis 216).
Grisebach (Fl. Rumel.) describes the anthers as eight, combined into four pha-
langes, as follows:—" Antherse 8 in phalangibas 4 distinctis petalo oppositis binatim
conjunctse;" and again, " Stamina tetradelpha per paria petalo opposita" (Grundriss
der Syst. Bot. 70), but in our specimens (and all others we have examined) the sta-
mens are as described by other authors, namely, four.opposite the petals, with some-
times a real or perhaps only apparent cohesion of the anthers. The anthers are com-
pletely enclosed within the middle lobes of the lateral petals daring impregnation,
when the lobes retain the $olUn in. contact with the stigma, exactly as the lateral
petals do in other Fumariacea* Indeed, the bud of Hypecoum scarcely differs from
that of a spurless Corydalis in appearance, the outer petals being sharply keeled at
their connivent apices, and the inner enclosing the anthers in their tips,
Bcrnhardi (Linnsea, viii.) regards the sepals as bracts, the two outer petals as
sepals, and the two inner as stamens and petals combined: but this appears to us
to be a purely hypothetical view, and not supported by any anatomical or morpho-
logical facts in tlie structure of these organs or their relations.
We have in vain endeavoured to find any character whereby to separate Cldavy-
spermum generically from Sypecoum; the articulation of the pod varies greatly ia
amount, and is shared by Hypecoum to a considerable extent. The forms of this
genus are, like all cut-leaved herbaceous annuals, excessively variable in habit and
foliage, and there are probably extremely few good species.
1. H. procumbens (Linn. Sp. PL 181); siliquis arcuatis sub-
compressis, sepalis ovatis, petalis exterioribus late obcuneato-trilobis,
interioribus alte trilobis, lobo intermedio spathulato margine subciliato.
—Sidthorp, Mor. Gnec. ii. 46. t. 155 ; SchkuJir, Handt>uch,*i. 90. t. 27;
DC. SysL ii. 102, Prodr. i. 123.
HAB. Panjab ad Peshawar, Vicary! Multan, EJgeworth! "Salt range/*
Fleming ! Beluchistan, Stocks / Afghanistan, Griffith !—(PL vere.) (t?. *.)
DISTRIB. Eegio Mediterranea 1 et Caspica! Asia Minor! Mesopo-
tamia ! Persia!
Herba procumbens, annaa, glauca, spithamsea. lolia radicalia in lacinias an-
gustas bitripinnatisecta, Cautes v. scapi 2-4, decumbentes v. ascendentes, teretes,
feeves. Sractea verticillataj, -^-1-pollicares, lacersa. Flares pauci, pedicellati; ala-
bastri nutantes, aurei, -| poll, lati, Sepala, fugacia, lato ovata v. ovato-lanceolata,
acuminata, petalis J- breviora. Petafa extcriora laic obovato- v. cuneato-rhombea,
plus minusve sinuato-triloba, lobis lateralibus lineari-pblongis obtusis, intermedio
spathulato, forma varia, oblonga, retusa v. bifida, margmibus fimbriato- v. sinuato-
dentatis. Anthera connective plus minusve ultra loculos producto ; filamenta dila-
tata. Stigma bicrure. Silica linearis, l-£~2£ poll, longa, 2 iin. lata, utrinquc at-
tenuata, compresso, spongiosa. Semina oblique obovata, arillo parvo instructa, testa